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(2003) Effects of habitat fragmentation on avian nesting success: a review of the evidence at multiple spatial scales. Biological Conservation 115(1):101–110. doi: 10.1016/S0006-3207(03)00098-3 54. Tews J, Brose U, Grimm V, Tielbörger K, Wichmann MC, Schwager M, et al. (2004) Animal species diversity driven by habitat heterogeneity/diversity: the importance of keystone structures. Journal of Biogeography 31(1):79–92. doi: 10.1046/j.0305-0270.2003.00994.x 55. Tozer CD, Nol E, Abraham KF (2010) Effects of local and landscape-scale habitat variables on abundance and