Morphometry of Fore Wing Venation for Identification of Net-Winged Insects of the Ukrainian Carpathians, with a Focus on Chrysopa (Neuroptera, Chrysopidae)

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A method to identify the net-winged insects without preparation of genitalia is proposed. The method is based on morphometric indices of fore wing venation and is applied to Neuroptera of the Ukrainian Carpathians. A key to nine neuropteran families is compiled, with an emphasis on Chrysopidae. Keys to eight genera and eleven species of the genus Chrysopa is provided. A statistical analysis of the variability of diagnostic traits is given for each species and possibility of using morphometric indices is substantiated for species identification.


  • Aspöck, H., Aspöck, U. and Hölzel, H. 1980. Die Neuropteren Europas: eine zusammenfassende Darstellung der Systematik. Ö kologie und Chorologie der Neuropteroidea (Megaloptera. Raphidioptera. Planipennia) Europas: Bd 1–2. Goecke & Evers, Krefeld.

  • Aspöck, H., Hölzel, H. and Aspöck, U. 2001. Kommentierter Katalog der Neuropterida (Insecta: Raphidioptera. Megaloptera. Neuroptera) der Westpaläarktis. Denisia. 2, 1–606. URL:

  • Aspöck, U. 1993. Geklärtes und Ungeklärtes im System der Neuroptera (Insecta: Holometabola). Mitteilungen der Deutschen Gesellschaft für Allgemeine und Angewandte Entomologie, 8, 451–456.

  • Aspöck, U. and Aspöck, H. 2007.‘Verbliebene Vielfalt vergangener Blüte. Zur Evolution. Phylogenie und Biodiversität der Neuropterida (Insecta: Endopterygota). Denisia, 20, 451–516. URL:

  • Brooks, S. J. 1997 An overview of the current status of Chrysopidae (Neuroptera) systematics. Deutsche Entomologische Zeitschrift. 44 (2), 267–275.

  • Dong, M., Xu, H., Wang, Y., Jia, C., Liu, Z. 2016. Revision of the genus Heterosmylus Krüger, 1913 from China (Neuroptera. Osmylidae). ZooKeys, 637, 107–128. URL:

  • Hölzel, H. 1984. Chrysopidae of the Palearctic Region — a review’. In: Gepps, J., Aspöck, H. and Hölzel, H., eds. Progress in World’s Neuropterology. Graz, 61–68. URL:

  • Breitkreuz, C. V., Winterton, S. L., Engel, M. S. 2015. Revision of the green lacewing subgenus Ankylopteryx (Sencera) (Neuroptera. Chrysopidae). ZooKeys, 543, 111–127. URL:

  • Letardi, A., Aspoeck, U. & Aspoeck., H. (2017). Fauna Europaea: Chrysopa. Chrysopidae. Neuroptera. Fauna Europaea version 2017.06.

  • Seredіuk, G. V. 2015. Green lacewings (Insecta: Neuroptera. Chrysopidae) fauna of Ukraine. Proceeding of the State Natural History Museum, 31, 141–148. URL: [In Ukrainian].

  • Serediuk, G. V. 2016. Neuropterida (Insecta. Neuroptera) of the Ukrainian Carpathians. Ukrainian Entomological Journal, 1–2 (11), 46–68.

  • Serediuk, G. V. 2017. General review of morphological characters in different taxa of net-winged insects (Insecta: Neuroptera) with new keys to identification on the basis of the fore wings’ structure. Kharkov Entomol. Soc. Gaz., 15 (1), 57–70.

  • Tauber, C. A. Sosa, F. 2015. Rediscovery of Nuvol umbrosus Navás (Neuroptera. Chrysopidae. Leucochrysini): a redescription and discussion. ZooKeys, 519, 141–153. URL:

  • Tauber, C. A., Sosa, F. Albuquerque, G. S. Tauber, M. J. 2017. Revision of the Neotropical green lacewing genus Ungla (Neuroptera. Chrysopidae). ZooKeys, 674, 1–188. URL:

  • Toschi, C. A. 1965. The taxonomy life histories and mating behavior of the green lacewings of Strawberry Canyon (Neuroptera, Chrysopidae). Hilgardia, 36, 391–433.

  • Winterton, S. L., Brooks, S. J. 2015. Review of the green lacewing genus Chrysacanthia Lacroix with a new species from Nigeria (Neuroptera. Chrysopidae). ZooKeys, 517, 71–81.

  • Zakharenko, A. V. 1993. The green lace-wings (Neuroptera. Chrysopidae) of Ukraine. The Kharkov Entomological Society Gazette. 1(1), 44–58. URL: [In Russian].

  • Zhao, Y., Yan, B., Liu, Z. 2015. A new species of the brown lacewing genus Zachobiella Banks from China (Neuroptera. Hemerobiidae) with a key to species. ZooKeys, 502, 27–37. URL:

Vestnik Zoologii

The Journal of National Academy of Sciences of Ukraine, Schmalhauzen Institute of Zoology

Journal Information

Cite Score 2016: 0.35

SCImago Journal Rank (SJR) 2016: 0.300
Source Normalized Impact per Paper (SNIP) 2016: 0.496


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