Review of the Ordovician stratigraphy and fauna of the Anarak Region in Central Iran

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The Ordovician sedimentary succession of the Pol-e Khavand area, situated on the northern margin of the Yazd block, has important differences from those in other parts of Central Iran. It has been established that the presumably terminal Cambrian to Lower Ordovician volcano-sedimentary Polekhavand Formation, exposed in the Pol-e Khavand area, has non-conformable contact with greenschists of the Doshakh Metamorphic Complex. The succeeding, mainly siliciclastic Chahgonbad Formation contains low to moderately diverse faunal assemblages, including brachiopods, cephalopods, trilobites and tentaculitids. The Darriwilian age of the lower part of the formation is well established by the co-occurrence of brachiopod genera Camerella, Phragmorthis, Tritoechia and Yangtzeella. The associated rich cephalopod fauna is different from the Darriwilian cephalopod associations of the Alborz terrane and may show some affinity with warm water faunas of North China and South Korea. It is likely that the Mid Ordovician fauna recovered from the lower part of the Chahgonbad Formation settled in the area sometime during a warming episode in the late Darriwilian. By contrast the low diversity mid Katian brachiopod association includes only three taxa, which occur together with the trilobite Vietnamia cf. teichmulleri and abundant, but poorly preserved tentaculitids questionably assigned to the genus Costatulites. This faunal association bears clear signatures linking it to the contemporaneous cold water faunas of the Arabian, Mediterranean and North African segments of Gondwana. Four brachiopod species recovered from the Chahgonbad Formation, including Hibernodonta lakhensis, Hindella prima, Lomatorthis? multilamellosa and Yangtzeella chupananica are new to science.

Alavi, M. 1991. Tectonic map of the Middle East: Tehran, Geological Survey of Iran, scale 1:5,000,000.

Andreeva, O.N. 1972. Brachiopods of the Ordovician Kuragan Formation of the South Urals. Paleontologicheskii Zhurnal, 1972 (1), 45-56. [In Russian]

Balashov, Z.G. 1968. Ordovician Endoceratoidea of the USSR, pp. 1-277. Izdatelstvo Leningradskogo Universiteta; Leningrad. [In Russian]

Balashova, E.A. 1966. Trilobites from the Ordovician and Silurian deposits of Pamir. Trudy Upravleniya Geologii Soveta Ministrov Tadzhikskoi SSR. Paleontologiya i Stratigraphiya , 2, 191-262. [In Russian]

Bassett, M.G., Popov, L.E. and Sokiran, E.V. 1999. Patterns of diversification in Ordovician cyrtomatodont rhynchonellate brachiopods. Acta Universitatis Carolinae, Geologica , 43, (1/2), 329-332.

Berberian, M. and King, G.C.P. 1981. Towards a paleogeography and tectonic evolution of Iran. Canadian Journal of Earth Science, 18, 210-265. DOI: 10.1139/e81-019

Berger, A.Ya. 1982. Ancient tentaculitides of Siberia. In Silurian of Siberian Platform, Sections, fauna and flora of the north-western part of the Tunguska Sineclise. Trudy Instituta Geologii i Geofiziki Sibirskogo Otdeleniya Akademii Nauk SSSR, 508, 132-141.

Billings, E. 1859. Description of a new genus of Brachiopoda and on the genus Cyrtodonta. Canadian Naturalist and Geologist, 4, 301-303.

Billings, E. 1862. New species of fossils from different parts of the lower, middle, and upper Silurian rocks of Canada. Palaeozoic Fossils, 1, 96-185.

Bogelopova, O.K., Kröger, B., Falahatgar, M. and Javidan, M. 2013. Middle Ordovician cephalopods from the Abarsaj area, northern Iran. In: Anders, L. and Mehlqvist, K. (Eds), Proceedings of the 3rd IGCP 591 Annual Meeting Lund, Sweden, 9-19 June 2013, Lund University, p. 50.

Boucot, A.J., Johnson, J.G. and Staton, R.D. 1964. On some atrypoid, retzioid, and athyridoid Brachiopoda. Journal of Paleontology, 38, 805-822. URL:

Burmeister, H. 1843. Die Organisation der Trilobiten aus ihren lebenden Verwandten entwickelt; nebst einer systematischen Uebersicht aller zeither beschriebenen Arten, pp. 1-147. Ray Society; Berlin.

Chen, J. 1976. Advances in the Ordovician stratigraphy of North China with a brief description of nautiloid fossils. Acta Palaeontologica Sinica, 15, 55-74.

Chen, J. and Zou, X.1984. Ordovician cephalopods of the Ordos area, China. Memoir of the Nanjing Institute of Geology and Palaeontology, 20, 1-84.

Cocks, L.R.M. 1979. A silicified brachiopod fauna from the Silurian of Iran. Bulletin of the British Museum (Natural History), Geology, 32, 25-42.

Cocks, L.R.M. and Rong, J. 2000. Order Strophomenida. In: Kaesler, R.L. (Ed.), Treatise on Invertebrate Paleontology, Vol. H, Brachiopoda 2 (revised), pp. 216-348. Geological Society of America and Kansas University Press; Boulder, Colorado and Lawrence, Kansas.

Cocks, L.R.M. and Fortey, R.A. 1988. Lower Palaeozoic facies and faunas around Gondwana. In: Audley-Charles, M. G. and Hallam, A. (Eds), Gondwana and Tethys. Geological Society, London, Special Publication, 37, 183-200. DOI: 10.1144/GSL.SP.1988.037.01.10

Colmenar, J. and Álvaro, J.J. 2014. Integrated brachiopodbased bioevents and sequence-stratigraphic framework for a Late Ordovician subpolar platform, eastern Anti-Atlas, Morocco. Geological Magazine, DOI:

Cooper, G.A. 1956. Chazyan and related brachiopods. Smithsonian Miscellaneous Collections, 27, 1-1245.

Cuvier, G.L.C.F.D. 1797. Tableau élémentaire de l’histoire des animaux, pp. i-xvi + 1-710 14 pl. Deterville; Paris.

Dalman, J.W. 1828. Uppställning och Beskrifning af de i Sverige funne Terebratuliter. Kongliga Svenska Vetenskapsakademien Handlingar, 3 (for 1827), 85-155.

Davidson, T. 1882. A Monograph of the British Fossil Brachiopoda. Part 5 (1). Devonian and Silurian Supplements, pp. 1-134. Monograph of the Palaeontographical Society; London.

Destombes, J. 1972. Les trilobites de la sous ordre des Phacopina de l’Ordovicien del L’Anti Atlas (Maroc). Notes et Mémoires du Service Géologique de Maroc, 240, 1-113.

Dewitz, H.H. 1880. Ueber einige ostprussiche Silurcephalopoden. Zeitschrift der Deutschen geologischen Gesellschaft, 32, 371-393.

Dürkoop, A. 1970. Brachiopoden aus dem Silur, Devon und Karbon in Afghanistan. Palaeontographica, series A, 134, 153-225.

Evans, D.H. 1994. Siphonal and cameral deposits in Allumettoceras and the position of the Tripteroceratidae (Nautiloidea). Paläontlogische Zeitshrift, 68, 321-337. DOI: 0.1007/BF02991346

Evans, D.H., Ghobadi Pour, M. and Popov, L.E. 2013. Review of the Early to Mid Ordovician orthoconic cephalopods from Iran. Bulletin of Geosciences, 88, 21-44. DOI: 10.3140.bull. geosci.1355.

Flower, R.H. 1946. Ordovician cephalopods of the Cincinnati region. Bulletin of American Paleontology, 29, 1-656.

Flower, R.H. 1962. Notes on the Michelinoceratida. New Mexico Bureau of Mines and Mineral Resources, Memoir, 10, 21-55.

Flower, R.H. 1964a. The nautiloid order Ellesmeroceratida (Cephalopoda). New Mexico Bureau of Mines and Mineral Resources, Memoir, 12, 1-234.

Flower, R.H. 1964b. Nautiloid shell morphology. New Mexico Bureau of Mines and Mineral Resources, Memoir, 13, 1-78.

Flower, R.H. and Kummel, B.1950. A classification of the Nautiloidea. Journal of Paleontology, 24, 604-616.

Foerste, A.F. 1926. Actinosiphonate, trochoceroid, and other cephalopods. Denison University Bulletin, Journal Science Laboratories, 1, 285-384.

Foord, A.H. 1887. On Endoceras duplexWahl., with descriptions of three new species of Endoceras from the Ordovician of Sweden and Russia contained in the British Museum. Annals and Magazine of Natural History, Series 5, 20, 393-409.

Fortey, R.A. and Cocks, L.R.M. 2003. Palaeontological evidence bearing on global Ordovician-Silurian continental reconstructions. Earth-Science Reviews, 61, 245-307. DOI: 10.1016/S0012-8252(02)00115-0

Ghavidel-Syooki, M., Popov, L.E., Ghobadi Pour, M., Álvaro, J.J. and Ehsani, M.H. 2015a. Late Ordovician and early Silurian brachiopods from the Zagros Ranges, Iran. Earth and Environmental Science Transactions of the Royal Society of Edinburgh, 105, 159-187. DOI:

Ghavidel-Syooki, M., Ghobadi Pour, M., Popov, L.E., Álvaro, J.J., Klishevich, I.A., Rakhmonov, U. and Ehsani, M.H. 2015b. Late Ordovician cephalopods, tentaculitides, machaeridians and echinoderms from Kuh-e Faraghan, High Zagros, Iran. Alcheringa, 39, ## [In press] DOI: 10.1080/03115518.2015.1052677

Ghobadi Pour, M. 2006. Early Ordovician (Tremadocian) trilobites from Simeh-Kuh, Eastern Alborz, Iran. In: M.G. Bassett and V.K. Deisler (Eds), Studies in Palaeozoic palaeontology. National Museum of Wales Geological Series, 25, 93-118.

Ghobadi Pour, M. and Popov, L.E. 2009. First report on the occurrence of Neseuretinus and Ovalocephalus in the Middle Ordovician of Iran. Acta Palaeontologica Polonica, 54, 125-133. DOI: 10.4202/app.2009.0113

Ghobadi Pour, M., Popov, L.E., Kebria-ee Zadeh, M.R. and Baars, C. 2011. Middle Ordovician (Darriwilian) brachiopods associated with the Neseuretus biofacies, eastern Alborz Mountains, Iran. Memoirs of the Association of Australasian Palaeontologists, 42, 263-283.

Hairapetian, V., Ghobadi Pour, M., Popov, L.E. and Modzalevskaya, T.L. 2012. Stegocornu and associated brachiopods from the Silurian (Llandovery) of Central Iran. Estonian Journal of Earth Sciences, 61, 82-104. DOI: 10.3176/earth.2012.2.02

Hairapetian, V., Ghobadi Pour, M., Popov, L.E., Hejazi, S.H., Holmer, L.E., Evans, D. and Sharafi, A. 2015. Ordovician of the Anarak Region: implications in understanding Early Palaeozoic history of Central Iran. Stratigraphy, 12, 22−30.

Hall, J. and Clarke, J.M. 1892−1895. An introduction to the study of the genera of Palaeozoic Brachiopoda. New York Geological Survey, Palaeontology, 8, (1), I-xvi+1-367, pl. 1-41; 8 (2), 318-394, pl. 21-84.

Harper, D.A.T., Mitchell, W.I., Owen, A.W. and Romano, M. 1985. Upper Ordovician brachiopods and trilobites from the Clashford House Formation, near Herbertstown, Co. Meath, Ireland. Bulletin of the British Museum (Natural History), Geology, 38, 287-308.

Hammann, W. and Leone, F. 1997. Trilobites of the post- Sardic (Upper Ordovician) sequence of southern Sardinia. Part 1. Beringeria, 20, 1-217.

Havlíček V. 1961. Rhynchonelloidea des böhmischenälteren Paläozoikums (Brachiopoda). Rozpravy Ústredního ústavu geologického, 27, 1-211.

Havlíček, V. 1971. Brachiopodes de l’Ordovicien du Maroc. Notes et Mémoires du Service Géologique du Maroc, 230, 1-135.

Havlíček, V. and Vanek, J. 1966. The biostratigraphy of the Ordovician of Bohemia. Sbornik Geologickych Ved, Paleontologie , 8, 7-68.

Hawle, I. and Corda, A.J.C. 1847. Prodrom einer monographie der Böhmischen Trilobiten. Abhandlungen der Bohmisch Gesellschaft Wissenschaft, 5, 117-292.

Holm, G. 1896. Om apikaläden hos Endoceras. Geologiska a Föreningens i Stockholm Förhandlingar, 18, 394-416.

Hupé, P. 1955. Classification des trilobites II. Annales de Paléontologie, 41, 91-304.

Hyatt, A. 1883-84. Genera of fossil cephalopods. Boston Society of Natural History, Proceedings, 22, p. 253-338. [253-272, December 1883; 273-338, January 1884.]

King, W. 1846. Remarks on certain genera belonging to the class Palliobranchiata. Annals and Magazine of Natural History, Series 1, 18, 26-42.

Kobayashi, T. 1927. Ordovician fossils from Corea and South Manchuria. Japanese Journal of Geology and Geography, 5, 173-212.

Kobayashi, T. 1931. Studies on the stratigraphy and palaeontology of the Cambro-Ordovician formation of Hua-lienchai and Niu-hsin-tai, south Manchuria. Japanese Journal of Geology and Geography, 8, 131-189.

Kobayashi, T. 1934. The Cambro-Ordovician formations and faunas of South Chosen. Palaeontology. Part I. Middle Ordovician faunas. Journal of the Faculty of Sciences, Imperial University of Tokyo, Section 2, 3, 329-519.

Kobayashi, T. 1935. Restudy of Manchuroceras with a brief note on the classification of Endoceroids. Journal of the Geological Society of Japan, 42, 436-452.

Kobayashi, T. 1937. Contributions to the study of the apical end of the Ordovician nautiloids. Japanese Journal of Geology and Geography, 14, 1-21.

Kolarova, F.N. 1925. The generic status of ‘Triplecia’ poloi. Bulletin of the Geological Society of China, 4, 215-219.

Kröger, B., Beresi, M.S. and Landing, E. 2007. Early orthoceratoid cephalopods from the Argentine Precordillera (Lower - Middle Ordovician). Journal of Paleontology, 81, 1263-1280. DOI: abs/10.1666/06-013.1

Kuhn, O. 1940. Palaozoologie in Tabellen, pp. 1-50. G. Fischer; Jena.

Kuhn, O. 1949. Lehrbuch der Paläozoologie, pp. i-v + 1-326. E. Schweizerbart; Stuttgart.

Lindenberg, H.G., Gröler, K., Jacobshagen, V. and Ibbeken, H. 1984. Post-Paleozoic stratigraphy, structure and orogenetic evolution of the southern Sabzevar zone and the Taknar block. Neues Jahrbuch für Geologie und Paläontologie, Abhhandlungen, 168, 287-326.

Ljashenko, G.P. 1955. New facts on the systematics of tentaculitids, nowakids and styliolinids. Bulleten Moskovskogo Obshchestva Ispytatelei Prirody, Otdelenie Geologichskoye, Novaia Seriya, 30, 94-95.

Malinky, J.M., Wilson, M.A., Holmer, L.E. and Lardeux, H. 2004. Chapter 22. Tube-shaped incertae sedis. In: Webby, B., Droser, M.L., Paris, F. and Percival, I. (Eds), The Great Ordovician Biodiversification Event. Columbia University Press, pp. 214-222.

Mansuy, H. 1908. Contribution a la Carte géologique de l’Indo-Chine: Paléontologie. Gouvernement Général de l’Indo-Chine, Direction Générale des Travaux Publics, Service des Mines, pp. 1-73. Imprimerie d’Extreˆme- Orien; Hanoi-Haiphong.

Martelli, A. 1901. Fossili del Siluriano Inferiore dellos Schensi (China). Societa Geologica Italiana Bollettino, 20, 295-310.

McCoy F. 1844. A synopsis of the characters of the Carboniferous Limestone fossils of Ireland pp. 1-274. Dublin University Press; Dublin.

Milne Edwards, H. 1840. Histoire naturelle des Crustacés, comprenant l’anatomie, la physiologie et la classification de ces animaux, 3, pp. 1-638. Roret; Paris.

Neuman, R.B. 1968. Paleogeographic implications of Or- dovician shelly fossils in the Magog belt of the northern Appalachian region. In: Zen, A, White, W.S., Hadley, J.B. and Thompson, J.B., Jr. (Eds), Studies of Appalachian Geology: Northern and Maritime pp. 35-48. Interscience Publishers (a division of John Wiley and Sons).

Percival, I.G., Wright, A., Nicoll, R S. and M.A. Hamedi 2009. Martellia and associated Middle Ordovician brachiopods from the Katkoyeh Formation, east-central Iran. Memoirs of the Association of Australasian Palaeontologists , 37, 315-325.

Nikitina, O.I., Popov, L.E., Neuman, R.B., Bassett, M.G. and Holmer, L.E. 2006. Mid Ordovician (Darriwilian) brachiopods of South Kazakhstan. In: Bassett, M.G. and V.K. Deisler, (Eds), Studies in Palaeozoic palaeontology. National Museum of Wales Geological Series, 25, 145-222.

Öpik, A. 1934. Über Klitamboniten. Tartu Universitatis (Dorpatensis), Acta and Commentationes, series A26, 4-239.

Pander, G.H. 1830. Beiträge zur Geognosie des Russichen Reiches, pp. 1-165. St. Petersburg,

Popov, L.E., Egerquist, E. and Zuykov, M.A. 2005. Ordovician (Arenig-Caradoc) Syntrophiidine brachiopods from the East Baltic region. Palaeontology, 48, 739-761. DOI: 10.1111/j.1475-4983.2005.00487.x

Popov, L.E., Bassett, M.G., Zhemchuzhnikov, V.G., Holmer, L.E. and Klishevich, I.A. 2009. Gondwanan faunal signatures from early Palaeozoic terranes of Kazakhstan and Central Asia: evidence and tectonic implications. In: Bassett, M.G. (Ed.) Early Palaeozoic Peri-Gondwanan Terranes: New Insights from Tectonics and Biogeography. The Geological Society, London, Special Publications, 325, 23-64. DOI:10.1144/SP325.3

Popov, L.E., Nikitin, I.F. and Sokiran, E.V. 1999. The earliest atrypides and athyridides (Brachiopoda) from the Ordovician of Kazakhstan. Palaeontology, 42, 625-661. DOI: 10.1111/1475-4983.00089

Popov, L. E., Vinn, O. and Nikitina, O.I. 2001. Brachiopods of the redefined family Tritoechiidae from the Ordovician of Kazakhstan and South Urals. Geobios, 32, 131-155. DOI: 10.1016/S0016-6995(01)80057-7

Popov, L., Hairapetian, V., Ghobadi Pour, M., Buttler, C., Evans, D.H., Hejazi, S.H. and Jahangir, H. 2014. Llandovery fauna of Iran during the post-extinction recovery. The Third International Symposium of the International Geosciences Programme Project 589 (IGCP- 589): Development of the Asian Tethyan Realm: genesis, process and outcomes. Tehran, 19-26 October, 2014, Abstract volume and proceedings, 105-110.

Ramezani, J. and Tucker, R.D. 2003. The Saghand region, central Iran: U-Pb geochronology, petrogenesis and implications for Gondwana tectonics. American Journal of Science , 303, 622-665. DOI: 10.2475/ajs.303.7.622

Rong, J. and Cocks, L.R.M. 1994. True Strophomena and a revision of the classification of strophomenind and ‘stropheodontid’ brachiopods. Palaeontology, 37, 651-694.

Rong, J., Harper, D.A.T., Zhan, R., Huang, Y. and Cheng, J. 2005. Silicified rhynchonelliform brachiopods from the Kuniutan Formation (Darriwilian: Middle Ordovician), Guiyang, South China. Palaeontology, 48, 1211-1240. DOI: 10.1111/j.1475-4983.2005.00506.x

Ross, R.J. 1972. Fossils from the Ordovician bioherm at Meiklejohn Peak, Nevada. United States Geological Survey, Professional Paper, 685, 1-47.

Reed, F.R.C. 1906. The Lower Palaeozoic fossils of the northern Shan States, Burma. Memoirs of the Geological Survey of India, Palaeontologia Indica, New Series, 2, 1-154.

Rozman, Kh.S. 1969. Late Ordovician brachiopods of the Siberian Platform. Paleontologicheskii Zhurnal, 1969 (3), 86-108.

Rubel, M. 1961. Lower Ordovician brachiopods of the superfamilies Orthacea, Dalmanellacea, and Syntrophiacea of East Baltic. Eesti NSV Teaduste Akadeemia Geoloogia Instituudi, Uurimuse, 6, 141-226.

Ruttner, A., Nabavi, M. and Hajian, J. 1968. Geology of the Shirgesht area (Tabas area, East Iran). Reports of the Geological Survey of Iran, 4, 1-133.

Salter, J.W. 1864. A monograph of the British trilobites from the Cambrian, Silurian and Devonian formations. Part 1. Monograph of the Palaeontographical Society, 16, 1-80.

Schallreuter, R., Hinz-Schallreuter, I., Balini, M. and Ferretti, A. 2006. Late Ordovician Ostracoda from Iran and their significance for palaeogeographical reconstructions. Zeitschrift für Geologische Wissenschaften, 34, 293-345.

Schmidt, H. 1965. Neue Befunde an Paläozoischen Rhynchonellacea (Brachiopoda). Senckenbergiana Lethaea, 46, 1-25.

Schuchert, C. 1893. A classification of the Brachiopoda. American Geologist, 11, 141-167.

Schuchert, C. 1913. Class 2. Brachiopoda In: von Zittel, K.A. (Ed.), Text-book of Palaeontology, 1, (2nd ed.), pp. 355-420. MacMillan; London.

Schuchert, C. and Cooper, G.A. 1931. Synopsis of the brachiopod genera of the suborders Orthoidea and Pentameroidea, with notes on the Telotremata. American Journal of Science, 22, 241-251.

Schuchert, C. and Cooper, G.A. 1932. Brachiopod genera of the suborders Orthoidea and Pentameroidea. Memoirs of the Peabody Museum of Natural History, 4, 1-270.

Sharkovski, M., Susov, M. and Krivyakin, B. (Eds) 1984. Geology of the Anarak area (Central Iran), Explanatory text of the Anarak quadrangle map, 1:250.000, V/O Technoexport Report, 19, pp. 1-143. Gological Survey of Iran; Tehran.

Sheehan, P.M. 1977. Late Ordovician and earliest Silurian meristellid brachiopods in Scandinavia. Journal of Paleontology , 51, 23-43.

Sweet, W.C. 1964. Orthocerida. In: Moore R.C. (Ed.), Treatise on invertebrate paleontology. Part K. Mollusca 3, Cephalopoda. General features, Endoceratoidea, Actinoceratoidea, Nautiloidea, Bactritoidea, pp. 216-261. Geological Society of America and University of Kansas Press; Boulder, Colorado and Lawrence, Kansas.

Teichert, C. 1933. Der bau der actinoceroiden cephalopoden. Palaeontographica, Abt. A., 78, 111-230.

Teichert, C. 1964. Endocerida. In: Moore, R.C. (Ed.), Treatise on invertebrate paleontology. Part K. Mollusca 3, Cephalopoda. General features, Endoceratoidea, Actinoceratoidea, Nautiloidea, Bactritoidea, pp. 160-189. Geological Society of America and University of Kansas Press; Boulder, Colorado and Lawrence, Kansas.

Ulrich, E.O. and Cooper, G.A. 1936. New genera and species of Ozarkian and Canadian brachiopods. Journal of Paleontology , 10, 616-631.

Ulrich, E.O. and Cooper, G.A. 1942. New genera of Ordovician brachiopods. Journal of Paleontology, 16, 620-626.

Villas, E. 1985. Braquiópodos del Ordovícico medio y superior de las Cadenas Ibéricas Orientales. Memorias del Museo Paleontológico de la Universidad de Zaragoza, 1, 1-223.

Waagen, W. 1883. Salt-Range fossils, vol. I, Part 4, Productus Limestone fossils, Brachiopoda. Memoirs of the geological Survey of India, Palaeontologia Indica (series 13), 2, 391-546.

Wahlenberg, G. 1821. Petrificata Telluris Svecanae. Nova Acta Regiae Societatis Scientiarum Upsaliensis, 8, 1-116.

Weber, V.N. 1948. Silurian trilobites of the USSR. 1: Lower Silurian trilobites. Monografiya po paleontologii SSSR, 69, (1), 1-113. [In Russian with English summary]

Williams, A. 1951. Llandovery brachiopods from Wales with special reference to the Llandovery District. Quarterly Journal of Geological Society, London, 107, 85-136.

Williams, A . 1962. The Barr and Lower Ardmillan Series (Caradoc) of the Girvan District, south-west Ayrshire, with descriptions of the Brachiopoda. Memoir of the Geological Society of London, 3, 1-267.

Williams, A. 1963. The Caradocian brachiopod faunas of the Bala District, Merionethshire. Bulletin of the British Museum (Natural History), Geology, 8, 329-471.

Williams, A. and Curry, G.B. 1985. Lower Ordovician Brachiopoda from the Tourmakeady Limestone, County Mayo, Ireland. Bulletin of the British Museum (Natural History), Geology, 38, 183-269.

Wirth, E. 1936. Über “Clitambonites” giraldii Martelli und Yangtzeella poloi Martelli aus dem Ordoviz Chinas. Paläontologische Zeitschrift, 18, 292-302.

Woodward, S.P. 1852. A manual of the Mollusca; or rudimentary treatise of recent and fossil shells, pp. i-xvi+1-486. London.

Yoh, S.S. 1945. On the inner structure of a new species of Yangtzeella. Bulletin of the Geological Society of China, 24, 11-14.

Yun, C.S. 2011. Ordovician cephalopods from the Jigunsan Formation, Taebaek-Yeongwol, Korea. Journal of the Paleontological Society of Korea, 27, 149-259.

Zhan, R. and Jin, J. 2005a. Brachiopods from the Dashaba Formation (Middle Ordovician) of Sichuan Province, Southwest China. Special Papers in Palaeontology, 74, 1-63.

Zhan, R. and Jin, J. 2005b. Brachiopods from the Middle Ordovician Shihtzupu Formation of Yunnan Province, China. Acta Palaeontologica Polonica, 50, 365-393.

Zhan, R., Jin, J. and Li, G. 2007. The Saucrorthis fauna (Brachiopoda) and its implication for the Ordovician radiation in South China. Acta Palaeontologica Sinica, 46, 515-522.

Zhan, R., Jin, J. and Chen, P. 2010. Early-Mid Ordovician Yangtzeella (Syntrophiidina, Brachiopoda) and its evolutionary significance. Palaeontology, 53, 77-96. DOI: 10.1111/j.1475-4983.2009.00921.x

Zhan, R., Luan, X., Huang, B., Liang, Y., Wang, G. and Wang, Y. 2014. Darriwilian Saucrorthis Fauna: implications for the Great Ordovician Biodiversification Event (GOBE). Estonian Journal of Earth Sciences, 63, 23-328.

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